The Naduo Formation or Nadu Formation is a geologic formation in Guangxi, southern China. The Naduo Formation preserves fossils dating back to the Middle or Late Eocene epoch of the Paleogene period, including a relatively diverse fauna of mammals.

Paleoenvironment In 1999, Gengwu Liu and Rongyu Yang published a study of fossil pollen collected in the Naduo Formation, identifying a diverse flora of 160 species, in 121 genera and 62 families. Based on the pollen, they could determine that the paleoenvironment of the Naduo Formation was likely a lowland basin, covered by a mixed evergreen and deciduous broad-leaved forest, probably similar to the environments that still dominate southern China today.

Mammal fossils

Aegialodontia +Genus/familySpeciesNotes/affinitiesImagesEodesmatodonE. spanios

Artiodactyla Genus/familySpeciesNotes/affinitiesImagesAnthracothemaA. rubricaeAn anthracothere.AnthracokeryxA. birmanicusAn anthracothere. The fossil record of Anthracokeryx in the Naduo Formation is very poor, and species assignments are questionable. A. moriturus may also be a synonym of Anthracotherium crassum. The recognition of Microbunodon in the Naduo Formation in 2010 casts doubt on identification of Anthracokeryx, since the two genera are difficult to tell apart dentally.A. moriturusA. sp.Bothriodon?B. chyelingensisAn anthracothere.Choeropotamidae or Helohyidae gen. nov.A new (undescribed) genus and species of cheropotamid or helohyid.EntelodontidaeEntelodontidae indet.An indeterminate entelodont.GobiomeryxG. sp.A ruminant.HeothemaH. belliaAn anthracothere.H. mediaHuananothemaH. imparilicaAn anthracothere.IndomeryxI. cotteriA ruminant.MicrobunodonM. sp.An anthracothere.NotomeryxN. besensisA ruminant.N. majorSuidae gen. nov. Asp. nov.A new (undescribed) genus and species of suid.Suidae gen. nov. Bsp. nov.A new (undescribed) genus and species of suid.Tayassuidae gen. nov.sp. nov.A new (undescribed) genus and species of peccary.TragulidaeTragulidae indet.A ruminant.

Carnivora Genus/familySpeciesNotes/affinitiesImagesCephalogaleC. sp. nov.A hemicyonine ("dog bear").cf. Cephalogalecf. C. sp.A hemicyonine ("dog bear").GuangxicyonZhai Renjie, Russell L. Ciochon, Tong Yongsheng, Donald E. Savage, Michael Morlo, Patricia A. Holroyd, and Gregg F. Gunnell. (2003) An aberrant amphicyonid mammal from the latest Eocene of the Bose Basin, Guangxi, China. Acta Palaeontologica Polonica 48 (2), 293-300G. sinoamericanusAn amphicyonid ("bear-dog"). Previously treated under the nomen nudum "Guangxicynodon sinocaliforniae".Pachycynodon?P.? sp.An amphicynodontid.

Hyaenodontia Genus/familySpeciesNotes/affinitiesImagesPropterodon?P.? sp.A hyaenodontid.

Mesonychia Genus/familySpeciesNotes/affinitiesImagesGuilestesG. acaresA mesonychid.G. cf. acarescf. Harpagolestescf. H. sp.A mesonychid.

Perissodactyla Genus/familySpeciesNotes/affinitiesImagesCaenolophusC. sp.An amynodont.DeperetellaD. sp.A deperetellid.EomoropusE. cf. quadridentatusAn eomoropid.GuixiaG. simplexA rhinoceros.HuananodonH. huiAn amynodont."Metatelmatherium""M". cf. browniiA brontothere. "Metatelmatherium" brownii does not belong to the genus Metatelmatherium, and the poor fossil material makes it difficult to identify. It may be a synonym of the dubious brontothere Sivatitanops birmanicum.ParamynodonP. sp.An amynodont.

References