Research A study aiming to reconstruct the life cycle of the Early Cambrian brachiopods is published by Madison & Kuzmina (2019). A study on the anatomy of plectambonitoid specimens from the Middle Ordovician of Russia, and on its implications for the knowledge of life cycles of these brachiopods, is published online by Madison & Kuzmina (2019). A study on the petrographic and geochemical preservation of Ordovician dalmanelloid shells from the Lexington Formation of Kentucky, Sheguindah Shale of Ontario and the Stony Mountain Formation of Manitoba, aiming to test the hypothesis of paleo-latitudinal zonation of the shelly benthos, is published by Azmy & Jin (2019). A study on the phylogenetic relationships among strophomenoid brachiopods and on the biogeographical changes in the strophomenoids through time (focusing on the impact of the Late Ordovician mass extinction on the evolutionary history of strophomenoids) is published by Congreve, Krug & Patzkowsky (2019). A study on the internal structure of the shell of Semiplanella carinthica is published by Pakhnevich (2019), who names a new tribe Semiplanellini in the subfamily Gigantoproductinae. A study on the diet of species of Gigantoproductus from the Viséan of Derbyshire (United Kingdom), and on its implications for the knowledge of mechanisms that enabled those brachiopod species to reach large sizes, is published by Angiolini et al. (2019). A study on the relative importance of brachiopods and bivalves in the fossil assemblages from the Carboniferous Pennsylvanian Breathitt Formation of Kentucky is published by Hsieh, Bush & Bennington (2019). Description of a specimen of a Permian brachiopod species Spiriferella protodraschei bearing ventrally directed spiralia (lophophore-supporting, coiled internal structures), and a study on the morphology of these structures and their implications for the knowledge of the feeding system of this brachiopod, is published by Lee et al. (2019). A study on the evolution of the body size of brachiopods from the Late Permian to the Middle Triassic, as indicated by brachiopod specimens from South China, is published by Chen et al. (2019). A study on the biogeography of Pliensbachian brachiopods in western Tethys Ocean is published online by Vörös & Escarguel (2019). A study on changes in the body size of benthic marine brachiopods and bivalves from the Lusitanian Basin (Portugal) before the Toarcian oceanic anoxic event is published by Piazza et al. (2019). A study on the impact of the early Toarcian extinction event on fossil brachiopods and bivalves known from the Iberian Range (Spain) is published by Danise et al. (2019).
New taxa Name Novelty Status Authors Age Type locality Country Notes ImagesAegiria aptaSp. novValidHuang et al.Silurian (Rhuddanian)Zhangwan FormationAlaskorhynchusGen. et sp. novValidBaranov & BlodgettDevonian (Pragian)Soda Creek LimestoneA member of Rhynchonellida belonging to the family Eatoniidae. The type species is A. sodacreekensis.Angustothyris dagysiSp. novValidGuo, Chen & HarperMiddle Triassic (Anisian)Qingyan FormationAngustothyris qingyanensisSp. novValidGuo, Chen & HarperMiddle Triassic (Anisian)Qingyan FormationSubsequently transferred to the genus Qianothyris.Anidanthus parvimucronataNom. novValidHe & Shen in He et al.Permian (Changhsingian)Talung FormationA member of Productida belonging to the family Linoproductidae and the subfamily Anidanthinae; a replacement name for Anidanthus mucronata He & Shen in He et al. (2005).Anidanthus subquadratusSp. novValidHe, Shi & Shen in He et al.Permian (Changhsingian)A member of Productida belonging to the family Linoproductidae and the subfamily Anidanthinae.Aporthophyla priscaSp. novValidCocks & PopovEarly OrdovicianArcullina? enokianiSp. novValidLee & Shi in Lee et al.Permian (Kungurian)Kozhim Formation Kozhim Rudnik FormationA member of Spiriferida belonging to the family Spiriferellidae.AskerinaGen. et sp. novValidBaarliOrdovician (Hirnantian) and Silurian (Aeronian)SolvikA member of the family Atrypidae. The type species is A. cymbula. Announced in 2019; the final version of the article naming was published in 2021. Aulacothyris stevensiSp. novValidMacFarlanEarly Jurassic (Aratauran)Diamond Peak GroupA member of Terebratulida belonging to the family Zeilleriidae.Aulonotreta neptuniSp. novValidHolmer et al.Ordovician (Darriwilian )A member of Lingulida belonging to the family Aulonotretidae.Australocoelia boucotiSp. novValidDe Rezende, Machado & PoncianoDevonianAustriellula iordanaeSp. novValidGaetani in Grădinaru & GaetaniTriassicA member of Rhynchonellida belonging to the family Norellidae and the subfamily Norellinae.BellistrophiaGen. et comb. novValidHolmer et al.CambrianAtei Formation Meagher FormationA member of Kutorginida. The type species is "Nisusia" deissei Bell (1941); genus also includes "Nisusia" montanensis Bell (1941).BernousiaGen. et sp. novValidGourvennecPaleozoicTindouf BasinA member of Spiriferida. Genus includes new species B. brevedorsata.Biernatia rhapsodySp. novValidLavié, Serra & FeltesMiddle and Late OrdovicianLas Aguaditas Formation Las Chacritas FormationA member of Acrotretida belonging to the family Biernatidae.BorellithyrisGen. et sp. novValidDulaiMiocene (Tortonian)Sant'Agata Fossili FormationA member of Terebratulida belonging to the family Megathyrididae. The type species is B. gaetanii.Cathaysiorthis xichuanensisSp. novValidHuang et al.Silurian (Rhuddanian)Zhangwan FormationCaucasorhynchellaGen. novValidGuo, Chen & HarperMiddle Triassic (Anisian)Chonosteges cooperiSp. novValidTorres-Martínez et al.PermianPaso Hondo FormationCleiothyridina pfaffenbergensisSp. novValidMottequin et al.Carboniferous (Tournaisian)Gleitsch FormationA member of Athyridida belonging to the family Athyrididae."Clorinda" symmetricaSp. novValidCocksSilurian (Llandovery)Conotreta andinaSp. novValidLavié, Serra & FeltesOrdovician (Darriwilian)Las Aguaditas Formation Las Chacritas FormationA member of Acrotretida belonging to the family Acrotretidae.Coolinia diabolicaSp. novValidCocksSilurian (Llandovery)CordatiaGen. et sp. novValidBrock & Claybourn in Betts et al.Early CambrianA member of Paterinata possibly belonging to the family Paterinidae. The type species is C. erinae.Craniops popoviSp. novValidCocksSilurian (Llandovery)Crassispirifer galinaeSp. novValidManankovPermianA member of Spiriferida.Crassumbo germanicusSp. novValidMottequin et al.Carboniferous (Tournaisian)Gleitsch FormationA member of Spiriferida belonging to the family Crassumbidae.Cyrtia wrekinensisSp. novValidCocksSilurian (Llandovery)Cyrtina evanescensSp. novValidGourvennecPaleozoicTindouf BasinA member of Spiriferinida.Cyrtospirifer aouinetensisSp. novValidGourvennecPaleozoicTindouf BasinCyrtospirifer (Elasmospirifer)Subgen. et sp. novValidGourvennecPaleozoicTindouf BasinThe subgenus includes new species C. (E.) djebiletensis.Cyrtospirifer robardetiSp. novValidGourvennecPaleozoicTindouf BasinDictyothyropsis vogliSp. novValidBujtor & VörösEarly CretaceousDihelictera askeriensisSp. novValidBaarliOrdovician (Hirnantian) and Silurian (Aeronian)SolvikA member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.Dinobolus athelstaniSp. novValidCocksSilurian (Llandovery)Eleutherokomma djezairensisSp. novValidGourvennecPaleozoicTindouf BasinA member of Spiriferida.Eospirifer majesticusSp. novValidCocksSilurian (Llandovery)Erbotreta elankiiSp. novValidUshatinskaya & KorovnikovCambrianA member of Acrotretida.Eucalathis giulioiSp. novValidDulaiMiocene (Tortonian)Sant'Agata Fossili FormationA member of Terebratulida belonging to the family Chlidonophoridae.FarewellirhynchiaGen. et sp. novValidBaranov & BlodgettDevonian (Pragian)Soda Creek LimestoneA member of Rhynchonellida belonging to the subfamily Leiorhynchinae. The type species is F. kulkovi.Fascistropheodonta? wiltzensisSp. novValidJansenEarly DevonianWiltz FormationA member of the family Strophodontidae.FelsithyrisGen. et sp. novValidMottequin & WeyerCarboniferous (Mississippian)A member of Spiriferida. The type species is F. hercynica.GarciaalcaldiaGen. et sp. novValidBaranov & BlodgettDevonian (Pragian)Soda Creek LimestoneA member of Rhynchonellida belonging to the subfamily Glossinunilinae. The type species is G. alaskensis.GhorispiriferGen. et comb. novValidMottequin & BriceDevonian (Famennian)A Cyrtiopsinae. The type species is "Cyrtiopsis graciosa" chakhaensis Brice (1971) (raised to the rank of a separate species G. chakhaensis), genus also includes "Cyrtiopsis" lapparenti Brice (1971).Hedeinopsis africanaSp. novValidGourvennecPaleozoicTindouf BasinA member of Spiriferida.Howellella minsterleyensisSp. novValidCocksSilurian (Llandovery)IngentistrophiaGen. et comb. novValidJansenEarly DevonianKatzenelnbogen Formation Seifen FormationA member of Orthotetida belonging to the family Areostrophiidae. The type species is "Orthothethes" ingens Drevermann (1904).Jonesea oepikiSp. novValidCocksSilurian (Llandovery)JordanithyrisGen. et sp. novValidFeldman et al.Middle Jurassic (Bathonian and Callovian)Hamam MughanniyyaA member of Terebratulida. Genus includes new species J. ardainensis. Announced in 2019; the final version of the article naming it was published in 2021. Joviatrypa nakremiSp. novValidBaarliSilurian (Aeronian)SolvikA member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021. JudinicaGen. et 2 sp. et comb. novValidOlenevaDevonianA member of Spiriferida belonging to the family Adolfiidae. The type species is J. siratchoica; genus also includes new species J. pseudodeflexa, as well as J. pseudomultifida (Vandercammen, 1955), J. biverrucosa (Vandercammen, 1955) and J. solita (Ljaschenko, 1959).Koeveskallina bifurcataSp. novValidGuo, Chen & HarperMiddle Triassic (Anisian)Qingyan FormationKosoidea australisSp. novValidZabini et al.Ordovician (Hirnantian)Iapó Formation