Many non-avian dinosaurs were feathered. Direct evidence of feathers exists for the following species, listed in the order currently accepted evidence was first published. In all examples, the evidence described consists of feather impressions, except those genera inferred to have had feathers based on skeletal or chemical evidence, such as the presence of quill knobs (the anchor points for wing feathers on the forelimb) or a pygostyle (the fused vertebrae at the tail tip which often supports large feathers).
Archaeopteryx lithographica (1861; possibly avialan) Ostromia crassipes (1970; possibly avialan) Avimimus portentosus (inferred 1987: ulnar ridge)Kurzanov, S.M. (1987). "Avimimidae and the problem of the origin of birds." Transactions of the Joint Soviet-Mongolian Paleontological Expedition, 31: 5-92. [in Russian]Chiappe, L.M. and Witmer, L.M. (2002). Mesozoic Birds: Above the Heads of Dinosaurs. Berkeley: University of California Press, 536 pp. Sinosauropteryx prima (1996) Fulicopus lyellii, an ichnotaxon, possible squatting Dilophosaurus or similar (1996) Protarchaeopteryx robusta (1997)Ji, Q., and Ji, S. (1997). "A Chinese archaeopterygian, Protarchaeopteryx gen. nov." Geological Science and Technology (Di Zhi Ke Ji), 238: 38-41. Translated By Will Downs Bilby Research Center Northern Arizona University January 2001 GMV 2124 (1997) Caudipteryx zoui (1998) Rahonavis ostromi (inferred 1998: quill knobs; possibly avialan) (HTML abstract) Shuvuuia deserti (1999) Beipiaosaurus inexpectus (1999) Sinornithosaurus millenii (1999) Caudipteryx dongi (2000) Caudipteryx sp. (2000) Microraptor zhaoianus (2000) Nomingia gobiensis (inferred 2000: pygostyle) Psittacosaurus sp.? (2002) Scansoriopteryx heilmanni (2002; possibly avialan)Czerkas, S.A., and Yuan, C. (2002). "An arboreal maniraptoran from northeast China." Pp. 63-95 in Czerkas, S.J. (Ed.), Feathered Dinosaurs and the Origin of Flight. The Dinosaur Museum Journal 1. The Dinosaur Museum, Blanding, U.S.A. PDF abridged version IVPP V13476 (2003) Yixianosaurus longimanus (2003; possibly avialan) Dilong paradoxus (2004)
Pedopenna daohugouensis (2005; possibly avialan) Jinfengopteryx elegans (2005) Juravenator starki (2006) Sinocalliopteryx gigas (2007) Velociraptor mongoliensis (inferred 2007: quill knobs) Epidexipteryx hui (2008; possibly avialan) Similicaudipteryx yixianensis (inferred 2008: pygostyle)
Anchiornis huxleyi (2009; possibly avialan) Tianyulong confuciusi? (2009) Incisivosaurus sp. (2010) Concavenator corcovatus? (inferred 2010: quill knobs?)Cuesta, Elena; Ortega, Francisco; Sanz, José Luis (October 2015). "Ulnar bumps of Concavenator: Quill Knobs or Muscular scar? Myological Reconstruction of the forelimb of Concavenator corcoavtus (Lower Cretaceous, Las Hoyas, Spain)." ResearchGate. Xiaotingia zhengi (2011; possibly avialan) Yutyrannus huali (2012)
Sciurumimus albersdoerferi (2012) Ornithomimus edmontonicus (2012) Ningyuansaurus wangi (2012) Eosinopteryx brevipenna (2013; possibly avialan) Jianchangosaurus yixianensis (2013) Aurornis xui (2013; possibly avialan) Changyuraptor yangi (2014) Kulindadromeus zabaikalicus? (2014) Citipati osmolskae (inferred 2014: pygostyle) Conchoraptor gracilis (inferred 2014: pygostyle) Deinocheirus mirificus (inferred 2014: pygostyle) Yi qi (2015; possibly avialan) Ornithomimus sp. (2015) Zhenyuanlong suni (2015) Dakotaraptor steini (inferred 2015: quill knobs) Apatoraptor pennatus (inferred 2016: quill knobs) DIP-V-15103 (2016; possibly avialan) Jianianhualong tengi (2017) Serikornis sungei (2017; possibly avialan) Caihong juji (2018; possibly avialan) Xingtianosaurus ganqi (2019) Ambopteryx longibrachium (2019; possibly avialan) Wulong bohaiensis (2020) Chirostenotes pergracilis? (2020) Dineobellator notohesperus (inferred 2020: quill knobs) Daurlong wangi (2022) Sinosauropteryx lingyuanensis (2025) Huadanosaurus sinensis (2025) Changzhousaurus sinensis (2026) Norellraptor barsboldi (2026)
Note that the filamentous structures in some ornithischian dinosaurs (Psittacosaurus, Tianyulong and Kulindadromeus) and the pycnofibres found in some pterosaurs may or may not be homologous with the feathers of theropods.Czerkas, S.A., and Ji, Q. (2002). "A new rhamphorhynchoid with a headcrest and complex integumentary structures." Pp. 15-41 in: Czerkas, S.J. (Ed.). Feathered Dinosaurs and the Origin of Flight. Blanding, Utah: The Dinosaur Museum. .
References