La Selva Biological Station is a protected area encompassing 1,536 ha of low-land tropical rain forest in northeastern Costa Rica. It is owned and operated by the Organization for Tropical Studies,Matlock, R., & Hartshorn, G. (1999). La selva biological station (OTS). Bulletin of the Ecological Society of America, 188-193. Retrieved from https://www.jstor.org/stable/20168346 a consortium of universities and research institutions from the United States, Costa Rica, and Puerto Rico.Moorman, R. (2006). Benefits of local residents visiting La Selva Biological Station, Costa Rica. Environmental Conservation, 88-99 Recognized internationally as one of the most productive field stations in the world for tropical forest research and peer-reviewed publications, La Selva hosts approximately 300 scientists and 100 university courses every year.Michener, W., Bildstein, K., McKee, A., Parmenter, R., Hargrove, W., McClearn, D., & Stromberg, M. (2009). Biological field stations: research legacies and sites for serendipity. BioScience, 300-310. The primary goal of La Selva Biological Station is to preserve and protect an intact forest, as well as providing laboratory facilities for tropical research and education. The research potential of the area is not only vital to tropical ecology, but it is also an important location in the effort to study relations between local communities and protected areas. In addition, its high diversity and ease of access to the Puerto Viejo-Horquetas highway makes La Selva an important ecotourism destination and environmental education center for tourists and the local community.
History
Leslie Holdridge established La Selva in 1953. Holdridge was an American botanist and climatologist who is known for his widely used classification system of land areas called The Holdridge Life Zones System. He originally purchased the land to use as a farm for experimenting with mixed agriculture, and was especially interested in experimenting with tree crops that could be planted without total clearing of the native forest.McDade, L., Bawa, K., Hespenheide, H., & Hartshorn, G. (1994). La selva: Ecology and natural history of a neotropical rain forest. Chicago: The University of Chicago Press In the beginning, La Selva was a challenge to get to; the trip required a long, treacherous dirt road followed by a 4-kilometer dugout canoe trip. In 1968 the Organization of Tropical Studies (OTS) bought the area for fifty thousand US dollars to be used as a private reserve and biological station. OTS was a small, 5-year-old organization created with the goal of facilitating research and education in the tropics. Holdridge encouraged OTS to use La Selva as a research site even before it became OTS property. Holdridge supported research at La Selva until his death in 1999. Since the station's beginnings in 1953, there have been immense changes: access is easy, buildings have been constructed, and the preserve has tripled in size. It is now one of the most important sites for tropical research in the world. In the past few decades, however, the human population around the Station have increased rapidly due to several factors, including the expansion of banana production and government settlement projects.
Location La Selva Biological Station is located in the Central Conservation Area (91,000 ha) in the north-eastern lowlands of Costa Rica (10°26´ N, 83°59´ W). It encompasses 1,536 ha of classic-lowland tropical rain forest. The Sarapiquí and Puerto Viejo Rivers border the Station to the north, the Peje River to the west, and the Sábalo-Esquina creeks to the east. Braulio Carrillo National Park (47,000 ha) is adjacent to the southern border of the Station and is connected via a 4–6 km wide protected corridor. The closest communities to the Station are La Flamínea (at its north-eastern border) and El Tigre (7 km south-east). Each of these communities has approximately 500 residents and they were established as recently as 1985. Puerto Viejo, the capital of the Cantón Sarapiquí in the province of Heredia, is the nearest major town, with approximately 1,163 residents, and is located seven kilometers north-east of La Selva. La Selva Biological Station is only 2 hours by car from San José airport.
Environment Of the total La Selva property (1,536 ha), 55% is characterized as species-rich, multilayered communities of primary forest. These forests contain impressive trees, lianas, epiphytes, palms, and many other broad-leaved monocots. The remaining areas of the reserve consist of abandoned pastures and plantations in various stages of succession, selectively logged secondary forest, or plots designated for experimental use. The reserve is located at the physiographic transition from the low, steep foothills of the Central Volcanic Cordillera to the vast Caribbean coastal plain in northeastern Costa Rica. At the northern end of the property the elevation is about 35 meters above sea level, but it quickly gives way to steep hills that reach up to 137 meters elevation at the southwest corner. The altitudinal transect connecting La Selva Biological Station to the main body of Braulio Carrillo National Park (approximately 55 km) extends from La Selva to Barva Volcano and includes tropical wet, premontane rain, lower montane rain, and montane rain forest ecosystems. Located at the confluence of the Puerto Viejo and Sarapiquí Rivers, which meet at the station's northern border, the preserve is surrounded on three sides by the natural barriers created by these rivers and their tributaries. The soils along the rivers are composed of Holocene and Pleistocene alluvia and consist primarily of inceptisols with occasional entisols. La Selva's uplands, on the other hand, begin several hundred meters inland from the two rivers and are composed primarily of utisols.
Temperature variation is slight year round, and daily temperatures can fluctuate from . As the climate is tropical wet, rain is common throughout the year, and the average annual rainfall is 4 meters (13 feet).La selva biological station. (n.d.). Retrieved from http://www.threepaths.co.cr/laselva_general_info.shtml The rainiest months are July, November, and December, while the least rainy season is from February to April. The dry season is rarely long or severe.
Flora and fauna
La Selva's biological resources are rich and largely intact which generates high biological diversity and productivity. The reserve is home to a wide range of wildlife, including large predators, rare birds, and an astonishing number of invertebrates. It is estimated that La Selva contains over 500,000 species, with more than half (about 300,000) consisting of insect species.Isenhower, J., & Cipollino (2009, June 15). Insect and spider biodiversity at La Selva biological station. Retrieved from http://facultyweb.berry.edu/mcipollini/Final Presentations/Insect and Spider Biodiversity.htm
Plants
Four major tropical life zones are protected between the La Selva and its connection to Braulio Carrillo National Park. Within this reserve, more than 5,000 species of vascular plants can be found, of which more than 700 species are trees.Organization for tropical studies. (n.d.). Retrieved from http://www.ots.ac.cr/index.php?option=com_content&task=view&id=162&Itemid=348 La Selva hosts a large variety of epiphytes, epiphylls, and many climbing aroids. Other common tree species include Welfia regia, Socratea exorrhiza, and most notably Pentaclethra macroloba, which occur in an unusual abundance.
Mammals
There are many mammalian species including large predators such as jaguars (Jaguar) and pumas (Puma (genus)). According to the OTS Website , five of the six species of felines in the country are found within the reserve. Some other species include the collared peccary (Tayassu tajacu) and Hoffmann's two-toed sloth (Choloepus hoffmanni).
Birds
The large number of bird species found here exceeds any other site in Central America. The most recent list published includes 467 species of birds.Zook, J., Alvarado, J., Rodolgo, A., Vargas, O., & Castro, E. (2012). Check-list of the birds of la selva. Retrieved from http://www.ots.ac.cr/images/downloads/biological-stations/la-selva/species-lists/bird-counting/birdsofls.pdf When taking into account the total number of species found one must understand that many are rare or may have been seen by mistake. Of these species, more than half breed locally, and many are altitudinal migrants or North American overwintering migrants.
Several ecological variables influence the vulnerability to habitat fragmentation, and other changes in the environment, in various groups of avifauna: diet, habitat, propensity to join mixed-species flocks, and nest type. Insectivorous birds are particularly sensitive to fragmentation, perhaps due to their dietary or foraging substrate specialization; forest under-story birds are most vulnerable to changes in their habitat; mixed-species flocks are especially sensitive to fragmentation as they typically roam over large areas; and certain nest types are more vulnerable to predation than others. According to Matlock and Hartshorn, the avifauna most sensitive to environmental changes, in general, are the mixed flocks of ant followers and under-story insectivores.
Along altitudinal gradients such as those observed in La Selva, species diversity and community composition of birds change rapidly. Elevational and latitudinal migrants, single and mixed flocks, threatened and endangered species, along with many other groups can be found in this area. Wide arrays of trophic groups are present, but the most common types are arboreal frugivores, arboreal frugivore/insectivores, foliage insectivores, and nectarivore/insectivores.
Other animals
La Selva's rivers contain an order of magnitude fewer species than the large, continental, Neotropical rivers such as the Amazon and Orinoco. Despite this notable difference, the two rivers and 13 streams flowing through La Selva support many species of Characidae, Poeciliidae, Cichlidae, and others. Large predators are also represented by the bushmaster Lachesis (genus). Despite the high diversity of plants and vertebrates, these numbers are dwarfed by its invertebrate fauna. Over 450 ant species have been collected at La Selva, with many still undescribed and there are more than 5,000 moth species (many undescribed).
A more complete species list can be found at the Organization for Tropical Studies site.
Research
There is a great amount of research conducted at La Selva Biological Station. Each year, more than 240 papers are published from studies performed there. The station has grown and expanded its facilities to include two laboratories, workspaces, an academic center, a workers’ lounge, a researchers’ lounge, a river station, an herbarium, and of course, the surrounding reserve all for research purposes. Researchers can live on site at the station for anywhere between a few days to several months. To accommodate researchers there are cabin dormitories, family housing, a laundry room, a dining hall, and even a gift shop.