2-Phospho-L-lactate guanylyltransferase (, CofC, MJ0887) is an enzyme with systematic name GTP:2-phospho-L-lactate guanylyltransferase. This enzyme catalyses the following chemical reaction
The enzyme characterised from methanogenic archaea and Paraburkholderia rhizoxinica combines 2-phospho-L-lactic acid and guanosine triphosphate to give L-lactyl-2-diphospho-5'-guanosine, with pyrophosphate (PPi) as a byproduct. The reaction is part of the biosynthesis of coenzyme F420.
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